Introduction

Salt stress increases ionic influxes, oxidant imbalances, cell-division impairment, and membrane degeneration（Munns and Tester, 2008; Gill and Tuteja, 2010; Shabala and Munns, 2012）

Plants respond to salt stress conditions by activating different signaling pathway, which assist plants in adjusting their metabolism to maintain a defensive regime（Khan et al., 2013a, 2015）

Plant growth-promoting endophytic bacteria（PGPEB）form symptomless colonies in healthy host plant tissue. They are generally considered the most effective class of microorganism in assisting host plants to cope with abiotic and biotic stresses（Khan et al., 2013a; Mandyan and Jumpponen, 2014）